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Ianthe thought she was so fucking smart when she achieved lyctorhood before anyone else without seeing any of the chambers. Thought she was so clever to just eat her cavalier whole against the will of everyone.
How stupid must she have felt when she watched the birth of Paul, and realized that if she'd done more work, tried just a little harder, taken a step back and slowed down she could have united hers and Corona's souls for eternity.
Corona who cried that she hadn't been chosen at her sister's ascension. Ianthe who hadn't chosen her because she couldn't bear to lose her, like a naive fool. Choosing immortality condemned Ianthe to inevitably lose Corona. That moment she started the stop watch, and I'll bet never even realized until that impromptu birthday party.
And to rub it in Paul tells her there is still time for her and Naberius to be perfect, like she cared a scrap about Naberius. but there is no time for her and Corona, she and Corona will never have that chance again.
God she must have screamed and screamed